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Exercise Physiology 1: General Information
This page provides basic exercise physiology information. The skater who wishes much more
extensive coverage of the subject is referred to Stephen Seiler's Site.
Athletic Performance Model
The first figure below shows Seiler's model of athletic performance and velocity in particular. This figure indicates the areas
which contribute to your speed. I will discuss the broader issues and not get into details like mitochondria.
Energy Systems
Where does your body get the energy it needs for racing? Three energy systems are discussed:
Aerobic, Anaerobic, and ATP-CP.
Aerobic
Aerobic conditioning and Max VO2 training:
Aerobic conditioning is not the same as increasing Max VO2 (VO2MAX is discussed in detail at the bottom of this page.
At this point consider it as the most oxygen your body can utilize) . The bulk
of your training is for aerobic conditioning. This is low intensity training
that will accomplish various physiological adaptations crucial for performance improvement.
These adaptations are
- Increase liver and muscle carbohydrate stores.
- Improved cardiac function (blood pumping system)
- Improved thermoregulatory system (helps keep you cool)
- Improved neuromuscular function (smooth skating)
- Improved fat burning capabilities (calories from fat spares carbohydrate)
- Increased number of mitochondria (energy producing structures)
- Increased number of capillaries around muscle fibers (better oxygen delivery and waste removal)
Max VO2 training will occur in the Race Phase and be accomplished by hard efforts above the
Anaerobic Threshold. Exercise at this level increases the amount of oxygen you can consume
during all out efforts. The greater the amount of oxygen consumed, the greater potential for
improved performance.
Anaerobic
Anaerobic metabolism or glycolysis occurs when the level of muscular activity causes the demand for
oxygen to exceed the body's ability provide it. In the absents of oxygen, the body will produce
energy but at a cost. The waste product of anaerobic metabolism is lactic acid. The more intense
the exercise, the faster that lactic acid is accumulated in the muscles and blood. Anaerobic
metabolism is responsible for the majority of energy production in all-out efforts lasting
approximately 1 to 10 minutes. The point at which lactic acid begins to quickly accumulate
in the blood is known as the anaerobic threshold or Lactic Threshold (LT).
A critical point needs to be made regarding the anaerobic threshold. The closer you can get your
anaerobic threshold to your Max VO2, the faster you will become. Simply having a high Max VO2
does not necessarily make you fast. Being able to sustain an effort close to your Max VO2 is what
will determine whether you are fast or not.
Lactic acid interferes with the muscles ability to contract. The burning sensation
experienced with the accumulation of lactic acid is the result of a change in muscular acidity.
The ability to tolerate and buffer the effects of this changing acidic environment is what anaerobic
training is all about.
ATP-CP
ATP (adenosine triphosphate) and CP (creatine phosphate) are the substances used for cellular
activity in all tissues, including muscle. This is the energy stored in muscles and available
immediately for use by the muscles in contraction and relaxation. A limited amount of ATP can
be stored in muscles. When the intensity of effort is maximal, this energy source is depleted in
20-30 seconds. After this time, intensity must decline or the effort must stop. ATP must be continually
replenished during physical activity. This energy system would come into play
with climbing a short, steep hill or sprinting. Training this energy system requires very short bouts
of all-out effort (10-15 sec) with long and probably complete recovery between efforts. Care must be
taken not to stimulate the anaerobic energy system with it's production of lactic acid. The objective
of training this system is to deplete the ATP and CP stores thus stimulating a compensatory
increase in ATP and CP stores. Additional neuromuscular conditioning causes this system to
respond quicker. Creatine Monohydrate supplements (which yield CP) are commonly available in health
food stores now. ATP supplements can also be obtained but the efficacy is debateable.
The figure on the left above shows how different physical activities utilize the three energy mechanisms. The figure on
the right shows how an Olympic cyclist was able to modify his lactate threshold and drop his race time
by 4 minutes through training and aerodynamic changes.
Lactic Acid
Lactic acid buildup ( acidosis) can cause burning pain,
especially in untrained muscles. Lactic acid accumulation can lead to
muscle exhaustion withing seconds if the blood cannot clear it away. A
strategy for dealing with lactic acid buildup is to relax the muscles at
every opportunity, so that the circulating blood can carry the lactic acid
away and bring oxygen to support aerobic metabolism. ...much of the lactic
acid is routed to the liver, where it is converted to glucose. A little
lactic acid remains in muscle tissue, where it is completely oxidized when
the oxygen supply is once again sufficient.
Heart Rate vs. Age
The above figure shows the safe (green), caution (yellow), and danger (red)heartbeat rate zones
for a typical person of the age on the bottom scale. A stopwatch or heartrate monitor can be used to determine your
exercise heart rate.
Your maximum heart rate is very useful because your VO2MAX is correlated with the heart rate. as shown above.
Consequently you can use a stopwatch or heartrate monitor to observe increases (or decreases) in
your VO2MAX. To get an absolute VO2MAX you will need specialized medical/sports equipment but you can
get a relative reading using the heartrate method with a stopwatch.
VO2 Max Defined
VO2 max is the maximum volume of oxygen consumed by the body each minute
during exercise, while breathing air at sea level. Because oxygen
consumption is linearly related to energy expenditure, when we measure
oxygen consumption, we are indirectly measuring an individual's maximal
capacity to do work aerobically.
"What are the determinants of VO2 max?"
Every cell consumes oxygen in order to convert food energy to usable ATP
for cellular work. Muscle cells that are contracting have high demands
for ATP. So it follows that they will consume more oxygen during exercise.
The sum total of billions of cells throughout the body consuming oxygen,
and generating carbon dioxide, can be measured using
volume-measuring and oxygen-sensing equipment.
The muscle's capacity to use oxygen exceeds the heart's capacity for delivery.
Long term endurance training can result in a 300% increase
in muscle oxidative capacity, but only about a 15 to 25% increase in VO2 max.
VO2 max can be altered artificially by changing the oxygen concentration
in the air. VO2 max can be dissociated from skeletal muscle characteristics.
Heart stroke volume, in contrast, is linearly related to VO2 max. Training results
in an increase in stroke volume and
an increase in maximal cardiac output. Greater capacity for oxygen
delivery is the result. More muscle can be supplied with oxygen
simultaneously while still maintaining necessary blood pressure levels.
Heart performance dictates VO2 max.
In general, absolute VO2 max favors the large endurance
athlete, while relative VO2 tends to be higher in smaller athletes.
For comparison, the average maximal oxygen consumption of an untrained
male in his mid 30s is about 40-45 ml/min/kg, and
decreases with age. The same person who undergoes a regular endurance
exercise program might increase to 50-55 ml/min/kg. A champion male masters
runner age 50 will probably have a value of over 60 ml/min/kg.
An Olympic champion 10000 meter runner will probably have a value
approaching or over 80 ml/min/kg!
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